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Company Split DNS Woes?

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We are in the process of a company split, because of multiple unrelated reasons both companies are keeping the local AD domain name which is a .com lets call it jamfish.com.  Company 1 JamFish has external dns of JamFish.com as well, company 2 has external DNS of JamLee.com.  When I split the exchange environment I am thinking I will have a DNS problem sending from JamLee.com to JamFish.com because internal dns for JamLee.com is JamFish.com?  Did that make any sense?  I know I will be removing the accepted domains where appropriate, but I am thinking JamLee emails will get lost internally eventually being undeliverable because of the AD domain name.  Thoughts?

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