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Exchange 2013 Sending but not receiving.

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I have a standalone active directory server and another server that I want to dedicate to exchange. 

I have everything setup properly. I think..

I can send email's fine. But if I reply or send a new email to an account on the exchange server about a day later I get a failure message: 

The error that the other server returned was:451 4.7.0 Temporary server error. Please try again later. PRX2

I know my domain DNS is setup properly because if I install Small Business Server 2011 everything works fine. 

The only thing I can think of that is different is SBS comes with a DNS and DHCP server already installed. I am letting my router handle that currently. Is there an issue with that?

Thank you for your help.


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